1. Bhanthumnavin, K., 1965. A cytological study of polyploidy in the genus Fragaria. Ph.D. Thesis, University of Reading, England.
2. Bors, R., 2000. A streamlined synthetic octoploid system that emphasizes F. vesca as a bridge species. Ph.D. Thesis, University of Guelph, p. 272.
3. Bors, R. & J.A. Sullivan, 2005. Interspecific hybridization of Fragaria vesca subspecies with F. nilgerrensis, F. nubicola, F. pentaphylla and F. viridis. Accepted J Amer Soc Hort Sci.
4. Bumagima, S.I. & E. Mesaros, 1969. Evaluation of the results of hybridization of the strawberry species Fragaria vesca L., F. viridis Duch. and F. moschata Duch. by the set of hybrid seed and their viability. Genetika, Moskva 5(5): 32–38.
5. Evans, W.D., 1964. Cytogenetic investigations in the genus Fragaria. Ph.D. thesis, University of Reading, England.