1. Begon, M., Harper, J. L. &Townsend, C. R. — 1986.Ecology: Individuals, Populations and Communities. — Blackwell, Oxford.
2. Bornemissza, G. F. — 1976. The Australian dung beetle project 1965–1975. —Aust. Meat Res. Comm. Rev., 30, 1–30.
3. Bornemissza, G. F. — 1979. The Australian Dung Beetle Research Unit in Pretoria. —S. Afr. J. Sci., 75, 257–260.
4. Crawley, M. J. — 1987. What makes a community invasible? In: Colonisation, Succession and Stability. (A. J. Gray, M. J. Crawley &P. J. Edwards, eds), pp. 429–453. —Blackwell, Oxford.
5. Davis, A. L. V. — 1987. Geographic distribution of dung beetles [Coleoptera: Scarabaeidae] and their seasonal activity in the southwestern Cape Province. —J. S. Afr. Entomol. Soc., 50, 275–285.