1. ; Department of Biology, Faculty of Science; Osaka City University; Osaka 558 Japan
2. ; University of Marketing and Distribution Sciences; Gakuen-nishi-machi 3-1, Nishi-ku Kobe 673 Japan
3. Forestry and Forest Product Research Institute; P.O. Box 16 Tsukuba Norin Kenkyu Danchi-nai Ibaraki 305 Japan