1. School of Biosciences, University of Birmingham, Edgbaston, B15 2TT, Birmingham, UK
2. Institute of Microbiology and Infection, University of Birmingham, Edgbaston, B15 2TT, Birmingham, UK
3. National Physical Laboratory, Hampton Road, TW11 0LW, Teddington, UK
4. School of Pharmacy, University of Nottingham, University Park, NG7 2RD, Nottingham, UK