1. ; Graduate School of Environmental Sciences; University of Tsukuba; Tsukuba, Ibaraki 305 Japan
2. ; Division of Changing Earth and Agro-Environment; National Institute of Agro-Environmental Sciences; Tsukuba, Ibaraki 305 Japan
3. ; Division of Environmental Biology; National Institute for Environmental Studies; Tsukuba, Ibaraki 305 Japan
4. ; Department of Environmental Science and Resources, Faculty of Agriculture; Tokyo University of Agriculture and Technology; Fuchu Tokyo 183 Japan
5. ; Institute of Biological Sciences; University of Tsukuba; Tsukuba, Ibaraki 305 Japan