1. ; Graduate School of Environmental Sciences; University of Tsukuba; Tsukuba, Ibaraki 305 Japan
2. ; Department of Biology, Faculty of Science; Tokyo Metropolitan University; Hachiohji-shi Tokyo 192-03 Japan
3. ; Division of Plant Ecology; National Institute of Agro-Enivronmental Sciences; Tsukuba, Ibaraki 305 Japan
4. ; University of Tsukuba; Tsukuba, Ibaraki 305 Japan