1. Graduate School of Agriculture, Kyoto University, Kitashirakawa Oiwake-cho, Sakyo, Kyoto 606-8502, Japan
2. Field Science Education and Research Center, Kyoto University, Kitashirakawa Oiwake-cho, Sakyo, Kyoto 606-8502, Japan
3. Faculty of Agriculture, Kagoshima University, 1-21-24 Korimoto, Kagoshima 890-0065, Japan