1. Department of Primary Education, Tsuru University, 3– 8–1Tahara, Tsuru, Yamanashi 402-8555, Japan
2. Asian Natural Environmental Science Center, The University of Tokyo, 1– 1–8Midori-cho, Nishitokyo, Tokyo 188-0002, Japan
3. Field Science Center, Graduate School of Agricultural Science Forest Ecology, Tohoku University, 232–3 Aza-yomogida, Naruko Onsen, Osaki, Miyagi 989-6711, Japan
4. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo 113-8657, Japan
5. National Institute of Polar Research, 10–3, Midori-cho, Tachikawa-shi, Tokyo 190-8518, Japan