1. Divisions of Bio-Resources, Frontier Science Research Center, University of Miyazaki, Kiyotake 5200, Miyazaki, Miyazaki 889-1692, Japan
2. The University of Tokyo Chichibu Forest, Graduate School of Agriculture and Life Science, The University of Tokyo, Chichibu, Saitama 368-0034, Japan
3. Interdisciplinary Graduate School of Agriculture and Engineering, University of Miyazaki, Miyazaki 889-2192, Japan
4. Sugadaira Montane Research Center, University of Tsukuba, Sugadaira-kogen 1278–294, Ueda, Nagano 386-2204, Japan