1. The United Graduate School of Agricultural Sciences, Ehime University, Matsuyama 790-8566, Japan
2. Faculty of Agriculture, Kochi University, B200, Monobe, Nankoku 783-8502, Japan
3. Forestry and Forest Products Research Institute, Tsukuba 305-8687, Japan
4. Faculty of Agriculture, Ehime University, Matsuyama 790-8566, Japan