1. 0000 0001 0373 6302 grid.428986.9 College of Food Science and Technology Hainan University 570228 Haikou People’s Republic of China
2. 0000 0000 9735 6249 grid.413109.e Tianjin Industrial Microbiology Key Laboratory, College of Biotechnology Tianjin University of Science and Technology 300457 Tianjin People’s Republic of China