1. Boero F, Bouillon J, Piraino S. 1998. Heterochrony, generic distinction and phylogeny in the family Hydractiniidae (Hydrozoa: Cnidaria). Zoologische Verhandelingen, 323(4): 25–36
2. Bouillon J. 1978. Hydroméduses de l’archipel des Séchelles et du Mozambique. Revue de Zoologie Africaine (in French), 92: 118–172
3. Bouillon J, Boero F. 2000. Phylogeny and classification of Hydroidomedusae. Thalassia Salentina, 24: 1–296
4. Bouillon J, Gravili C, Pages F, et al. 2006. An Introduction to Hydrozoa. Vol 194. Paris: Publications Scientifiques du Museum Paris Paris France, 1–591
5. Edwards C. 1972. The hydroids and the medusae Podocoryne areolata, P. borealis and P. carnea. Journal of the Marine Biological Association of the United Kingdom, 52(1): 97–144, doi: https://doi.org/10.1017/S0025315400018609