1. Institute for Water Research, Rhodes University, Makhanda, South Africa
2. Freshwater Research Centre, Scarborough, South Africa
3. Faculty of Science, Nelson Mandela University, Gqeberha, South Africa
4. Department of Freshwater Invertebrates, Albany Museum, Makhanda, South Africa
5. Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa