1. Department of Ichthyology and Fisheries Science, Rhodes University, Makhanda, South Africa
2. DSI/NRF Research Chair in Inland Fisheries and Freshwater Ecology, South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, South Africa
3. Centre for Invasion Biology, SAIAB, Makhanda, South Africa
4. South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, South Africa
5. Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa