1. Ezemvelo KZN Wildlife, Pietermaritzburg, South Africa
2. Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa
3. Department of Biodiversity and Conservation Biology, University of the Western Cape, Cape Town, South Africa
4. School of Science and Engineering, University of the Sunshine Coast, Sippy Downs, Australia
5. Centre for African Conservation Ecology, Department of Zoology, Nelson Mandela University, Port Elizabeth, South Africa