1. Botany Department and Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa
2. DST/NRF Research Chair in Shallow Water Ecosystems, Nelson Mandela University, Port Elizabeth, South Africa
3. Department of Biochemistry and Microbiology, Rhodes University, Grahamstown, South Africa
4. South African Institute for Aquatic Biodiversity (SAIAB), Grahamstown, South Africa