1. Department of Conservation Ecology and Entomology, Stellenbosch University, Stellenbosch, South Africa
2. Department of Mathematical Sciences, Stellenbosch University, Stellenbosch, South Africa
3. African Institute for Mathematical Sciences (AIMS) South Africa, Cape Town, South Africa
4. The Southern African Foundation for the Conservation of Coastal Birds (SANCCOB), Cape Town, South Africa
5. Department of Biodiversity and Conservation Biology, University of the Western Cape, Cape Town, South Africa