1. Department of Biodiversity and Conservation Biology, University of the Western Cape, Cape Town, South Africa
2. Unit for Environmental Sciences and Management, School of Biological Sciences, North-West University, Potchefstroom, South Africa
3. South African Institute for Aquatic Biodiversity (SAIAB), Grahamstown, South Africa
4. Elwandle Coastal Node, South African Environmental Observation Network (SAEON), Port Elizabeth, South Africa