1. Institute for Coastal and Marine Research, Nelson Mandela University, Gqeberha, South Africa
2. Department of Biological Sciences, University of Cape Town, Cape Town, South Africa
3. Department of Biology, University of New Brunswick, Fredericton, Canada
4. Aquatic Ecosystem Services, Makhanda, South Africa
5. South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, South Africa