1. Zoology Department and the Institute for Coastal and Marine Research, Nelson Mandela University, Gqeberha (Port Elizabeth), South Africa
2. South African Environmental Observation Network (SAEON), Elwandle Coastal Node, Gqeberha, South Africa
3. Institute for Coastal and Marine Research, Nelson Mandela University, Gqeberha, South Africa
4. The Energy Institute, Colorado State University, Fort Collins, Colorado, United States
5. Department of Oceanography and the Institute for Coastal and Marine Research, Nelson Mandela University, Gqeberha, South Africa