1. Department of Plant and Soil Sciences, University of Pretoria, Pretoria, South Africa
2. South African Environmental Observation Network (SAEON): Arid Lands Node, Kimberley, South Africa
3. CapeNature, Assegaaibosch Nature Reserve, Jonkershoek, Stellenbosch, South Africa
4. Private, 7 St George Street, Somerset West, South Africa
5. Environment and Nature Conservation, Research and Development Support, Springbok, South Africa