1. Copernicus Institute for Sustainable Development, Utrecht University, Utrecht, The Netherlands
2. Soil Fertility & Analytical Services, KwaZulu-Natal Department of Agriculture & Rural Development, Pietermaritzburg, South Africa
3. School of Agricultural, Earth and Environmental Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa
4. Department of Wildlife, Fish and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden
5. Centre for African Conservation Ecology, Nelson Mandela University, Gqeberha, South Africa
6. South African Environmental Observation Network (SAEON), Grasslands, Forests and Wetlands Node, Pietermaritzburg, South Africa