1. School of Natural Resource Management, Nelson Mandela University, George, South Africa
2. REHABS International Research Laboratory, CNRS-Université Lyon 1-Nelson Mandela University, George, South Africa
3. Department of Geography and the Earth and Environmental Systems Institute, The Pennsylvania State University, Pennsylvania, United States of America
4. Eugène Marais Chair of Wildlife Management, Mammal Research Institute, University of Pretoria, Pretoria, South Africa