1. Sappi Shaw Research Centre, Sappi Forests, Howick, South Africa
2. Tree Protection Co-operative Programme (TPCP), Department of Plant and Soil Sciences, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, Pretoria, South Africa
3. Tree Protection Co-operative Programme (TPCP), Department of Biochemistry, Genetics and Microbiology, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, Pretoria, South Africa