1. South African National Biodiversity Institute (SANBI), Pretoria, South Africa
2. Department of Agriculture and Animal Health, University of South Africa, Johannesburg, South Africa
3. School of Health and Life Sciences, Teesside University, Middlesbrough, United Kingdom
4. National Institute for Theoretical and Computational Sciences (NITheCS), KwaZulu-Natal, South Africa
5. FitzPatrick Institute of African Ornithology, University of Cape Town, Cape Town, South Africa
6. Organisms and Environment Division, Cardiff University, Cardiff, United Kingdom