1. FitzPatrick Institute of African Ornithology, DSI/NRF Centre of Excellence, University of Cape Town, Cape Town, South Africa
2. Birds and Environmental Change Programme, Climate Change and BioAdaptation Division, South African National Biodiversity Institute, Cape Town, South Africa
3. Pacific Biodiversity Institute, University of Washington at Bothell, Anacortes, Washington, United States
4. Botany and Zoology Department, Stellenbosch University, Stellenbosch, South Africa