1. School of Biological Sciences, University of London, Egham, United Kingdom
2. Institute of Zoology, Zoological Society of London, London, United Kingdom
3. Biodiversity Inventory for Conservation, Glabbeek, Belgium
4. Both authors contributed equally to this manuscript
5. Department of Zoology, University of Cambridge, Cambridge, United Kingdom
6. FitzPatrick Institute of African Ornithology, University of Cape Town, Cape Town, South Africa
7. Maputo Natural History Museum, Maputo, Mozambique
8. Aquatic and Terrestrial Ecology, Royal Belgian Institute of Natural Sciences, Brussels, Belgium