1. Department of Biological Science, Florida State University, Tallahassee, Florida, USA
2. National Zoological Gardens of South Africa, Pretoria, South Africa
3. Department of Genetics, University of the Free State, Bloemfontein, South Africa
4. SARCHI Chair on Biodiversity Value and Change, School of Mathematical and Natural Sciences, University of Venda, Thohoyandou, South Africa