1. a CIRAD-ES, Unité de recherche Animal et Gestion Intégrée des Risques , TA 30/E Campus international de Baillarguet, 34398, Montpellier, France
2. b US Geological Survey Western Ecological Research Center , 505 Azuar Drive, Vallejo, CA, 94592, USA
3. c EMPRES Wildlife Unit, Animal Health Service, Animal Production and Health Division , Food and Agriculture Organization of the United Nations , Viale delle Terme di Caracalla, Rome, 00153, Italy
4. d Wetlands International , PO Box 471, 6700 AL, Wageningen, The Netherlands