1. a Department of Botany , University of Cape Town , Private Bag X3, Rondebosch, 7701, South Africa
2. b DST/NRF Centre of Excellence at the Percy FitzPatrick Institute of African Ornithology , Private Bag X3, Rondebosch, 7701, South Africa
3. c División Ornitología , Museo Argentino de Ciencias Naturales “B. Rivadavia” Av. Ángel Gallardo 470 (C1405DJR) , Buenos Aires, Argentina
4. d Department of Human Biology, Faculty of Health Sciences , University of Cape Town , Anzio Rd, Observatory, 7925, South Africa