1. a Department of Zoology , North Carolina State University , Campus Box 7617, Raleigh, NC, 27695-7617, USA
2. b School of Animal, Plant and Environmental Sciences , University of the Witwatersrand , Private Bag 3, Wits, 2050, South Africa
3. c School of Biological and Conservation Sciences , University of KwaZulu-Natal , Scottsville, 3209, South Africa
4. d Information Technology Division , Rhodes University , PO Box 94, Grahamstown, 6140, South Africa
5. e Department of Ornithology , National Museum , PO Box 266, Bloemfontein, 9300, South Africa
6. f PO Box 3076, Paulington, Mutare, Zimbabwe