1. a Animal Demography Unit, Department of Zoology , University of Cape Town , Rondebosch, 7701, South Africa
2. b Department of Zoology and Entomology , Rhodes University , PO Box 94, Grahamstown, 6140, South Africa
3. c Amicus Veterinary Centre , 90 Marshall Lake Road, Shirley, Solihull, UK
4. d Deceased