1. Department of Medicine, Ninewells Hospital and Medical School, University of Dundee, Dundee, Scotland
2. School of Life Sciences, Heriot Watt University, Edinburgh, Scotland
3. Division of Molecular Physiology, College of Life Sciences, University of Dundee, Dundee, Scotland
4. College of Life Sciences and Medicine, University of Aberdeen, Aberdeen, Scotland
5. Department of Pharmacology and Neurosciences, University of Dundee, Dundee, Scotland
6. School of Biomedical Sciences, Graduate Entry Medical School, University of Nottingham, Derby City Hospital, Derby, U.K