1. 1Division of Population Health and Genomics, School of Medicine, University of Dundee, Ninewells Hospital and Medical School, Dundee, U.K.
2. 2Division of Systems Medicine, School of Medicine, University of Dundee, Ninewells Hospital and Medical School, Dundee, U.K.
3. 3Usher Institute, College of Medicine and Veterinary Medicine, Edinburgh, U.K.