1. Laboratory of Molecular Genetics, Gunma University School of Medicine, Gunma University Maebashi, Gunma, Japan
2. Department of Cell Biology, and the Laboratory of Gene Analysis, Gunma University School of Medicine, Gunma University Maebashi, Gunma, Japan
3. Department of Molecular Medicine, Institute for Molecular and Cellular Regulation, Department of Laboratory Medicine, Gunma University School of Medicine, Gunma University Maebashi, Gunma, Japan
4. Howard Hughes Medical Institute, and the Departments of Biochemistry and Molecular Biology, and Medicine University of Chicago Chicago, Illinois