1. School of Biological Sciences, University of Wales Bangor, Bangor, Gwynedd LL57 2UW, Wales, United Kingdom (C.L., O.A.K., J.F.F., A.D.T.); and
2. Institute of Grassland and Environmental Research, Plas Gogerddan, Aberystwyth, Ceredigion SY 23 3EB, Wales, United Kingdom (J.G., C.J.P.)