1. Australian Research Council Centre of Excellence in Plant Energy Biology, Plant Research Centre, School of Agriculture, Food and Wine, Waite Research Institute, University of Adelaide, Waite Campus, Glen Osmond 5064, South Australia, Australia
2. Biochemistry and Plant Molecular Physiology, Université de Montpellier, Centre National de la Recherche Scientifique, Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement, Institut Agro, Montpellier 34060, France