1. College of Life Sciences, Qingdao University, Qingdao 266071, China
2. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
3. College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China
4. State Key Laboratory of Plant Physiology and Biochemistry, College of Biological Sciences, China Agricultural University, Beijing 100193, China