1. Allsopp, P.G. & Hutchinson, P.M. (2019) Extralimital Dynastinae (Coleoptera: Scarabaeidae) in Australia. Journal of Insect Biodiversity, 12, 48–77. https://doi.org/10.12976/jib/2019.12.2.2
2. Carne, P. B. (1957) Systematic revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp.
3. Cassis, G., Meades, L., Harris, R., Reid, C., Carter, G., Wilkie, L. & Jefferys, E. (2003) Lord Howe Island terrestrial invertebrate biodiversity and conservation. Report to the New South Wales National Parks and Wildlife Service by the Australian Museum Centre for Biodiversity and Conservation Research, Sydney, 88 pp.
4. Cumpston, D.M. (1940) On the external morphology and biology of Heteronychus sanctaehelenae Blanch. and Metanastes vulgivagus Olliff. Proceedings of the Linnean Society of New South Wales, 65, 289–300.
5. Darwin, C.R. (1859) On the origin of species by means of natural selection or, the preservation of favoured races in the struggle for life. John Murray, London, 502 pp. https://doi.org/10.5962/bhl.title.82303