1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, Washington 98105, USA
2. Western Ecological Research Center, Olympia Substation, US Geological Survey, Olympia, Washington 98516, USA
3. Northwest Fisheries Science Center, National Oceanic and Atmospheric Administration, Seattle, Washington 98112, USA
4. School of the Environment, Washington State University Vancouver, Vancouver, Washington 98686, USA
5. Graduate Program in Ecology, Evolution, Ecosystems, and Society, Dartmouth College, Hanover, New Hampshire 03755, USA
6. Western Fisheries Research Center, US Geological Survey, Seattle, Washington 98115, USA