1. Biological Station Rybachy, Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia
2. Bird Ringing and Scientific Information Centre of Russia, A.N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninsky Prospekt 33, 119071 Moscow, Russia
3. Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia; Saint Petersburg State University, Universitetskaya Emb. 7–9, 199034 Saint Petersburg, Russia