1. School of Life Sciences, Sun Yat-Sen University, Xingangxi Road, 135, 510275, Guangzhou, PR China
2. These authors contributed equally to this work.
3. Department of Microbiology, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China
4. Institute of Chinese Medicine, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China