1. *Center for Retrovirus Research, The Ohio State University, Columbus, OH
2. †Department of Veterinary Biosciences, The Ohio State University, Columbus, OH
3. ‡Molecular, Cellular, and Developmental Biology Program, The Ohio State University, Columbus, OH
4. §Viruses and Emerging Pathogens Program, Infectious Diseases Institute, The Ohio State University, Columbus, OH
5. ¶Department of Microbial Infection and Immunity, The Ohio State University, Columbus, OH