1. Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, 88 Xuefu Road, Kunming Yunnan 650223, China; Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical
Garden, Chinese Academy of Sciences, Menglun, Yunnan 666303, China; Graduate University of the Chinese Academy of Sciences, Beijing 100039, China
2. Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, 88 Xuefu Road, Kunming Yunnan 650223, China; Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden,
Chinese Academy of Sciences, Menglun, Yunnan 666303, China
3. Institute of Ecology and Geobotany, Yunnan University, Kunming 650091, China
4. Botany Department, Natural History Museum, London, SW7 5BD, U.K.; State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China