1. Biosciences Research Division, Department of Primary Industries, Bundoora, VIC 3083, Australia
2. Cooperative Research Centre for Sheep Industry Innovation, Armidale, NSW 2351, Australia
3. Faculty of Land and Environment, University of Melbourne, Parkville, VIC 3010, Australia
4. School of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia
5. La Trobe University, Bundoora, VIC 3086, Australia