1. Agricultural Research Council – Plant Health and Protection, P.O. Box 1055, Hilton, 3245, South Africa
2. Centre for Biological Control, Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa
3. Weed Research Unit, Invasive Species Biosecurity, NSW Department of Primary Industries, Biosecurity and Food Safety, 1447 Forest Road, Orange, NSW 2800, Australia
4. Department of Zoology and Entomology, Faculty of Natural and Agricultural Sciences, University of the Free State, P.O. Box 339, Bloemfontein 9300, South Africa