1. Centre for Biological Control, Botany Department, Rhodes University, Makhanda, 6140 South Africa
2. Agricultural Research Council – Plant Health and Protection, Private Bag X6006, Hilton, 3245 South Africa
3. Centre for Biological Control, Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa
4. CABI, Rues des Grillions 1, Delémont 2800, Switzerland