1. Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University , Nanjing 210037, Jiangsu , China
2. College of Life Science, Neijiang Normal University , Neijiang 641000, Sichuan , China
3. Key Laboratory of National Forestry and Grassland Administration on Subtropical Forest Biodiversity Conservation, College of Biology and the Environment, Nanjing Forestry University , Nanjing 210037 , China