1. Quantitative Biology Research Group, Exploratory Research Center on Life and Living Systems (ExCELLS), National Institutes of Natural Sciences , 5-1 Higashiyama, Myodaiji-cho, Okazak, Aichii, 444-8787 Japan
2. Division of Cellular Dynamics, National Institute for Basic Biology , Nishigonaka 38, Myodaiji, Okazaki, Aichi, 444-8585 Japan
3. Graduate School of Biostudies, Kyoto University , Kitashirakawa-Oiwakecho, Sakyo, Kyoto, 606-8502 Japan
4. Department of Applied Biological Science, Faculty of Science and Technology, Tokyo University of Science , 2641 Yamazaki, Noda, Chiba, 278-8510 Japan
5. Department of Basic Biology, SOKENDAI (The Graduate University for Advanced Studies) , Nishigonaka 38, Myodaiji, Okazaki, Aichi, 444-8585 Japan
6. Division of Quantitative Biology, National Institute for Basic Biology, National Institutes of Natural Sciences , 5-1 Higashiyama, Myodaiji-cho, Okazaki, Aichi, 444-8787 Japan
7. Department of Basic Biology, School of Life Science, SOKENDAI (The Graduate University for Advanced Studies) , 5-1 Higashiyama, Myodaiji-cho, Okazaki, Aichi, 444-8787 Japan