1. Ecosystem Dynamics and Forest Management Group , School of Life Sciences, , Hans-Carl-von-Carlowitz-Platz 2, 85354 Freising , Germany
2. Technical University of Munich , School of Life Sciences, , Hans-Carl-von-Carlowitz-Platz 2, 85354 Freising , Germany
3. Gund Institute for Environment , , 617 Main Street, Burlington, VT 05405 , United States
4. University of Vermont , , 617 Main Street, Burlington, VT 05405 , United States